As of November 7, 2026, 4 holders' latest Schedule 13D or 13G filings report owning 5% or more of Codexis, Inc. The largest reported stake is Aberdeen Group plc's, at 9.86%. 17 filings are on record since July 30, 2025 (Schedule 13G/A, October 30, 2026).
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Key facts
- · Reporting person: Opaleye Management Inc. · Sole voting: 0 · Shared voting: 2,590,000 · Sole dispositive: 0 · Shared dispositive: 2,590,000 · SCHEDULE 13G/A, 2026-11-06
- · Reporting person: Aberdeen Group plc · Sole voting: 0 · Shared voting: 10,797,702 · Sole dispositive: 0 · Shared dispositive: 10,797,702 · SCHEDULE 13G/A, 2026-10-30
- · Reporting person: Lytton Laurence W · Sole voting: 5,498,680 · Shared voting: 94,292 · Sole dispositive: 5,498,680 · Shared dispositive: 94,292 · SCHEDULE 13G/A, 2026-08-13
- · Reporting person: Telemark Asset Management, LLC · Sole voting: 0 · Shared voting: 7,846,411 · Sole dispositive: 0 · Shared dispositive: 7,846,411 · SCHEDULE 13G, 2026-07-29
- · Holder: Aberdeen Group plc · Percent: 7.45% · SCHEDULE 13G/A
Codexis, Inc. has 17 Schedule 13D or 13G filings on record since 2025-07-30. 4 holders' latest filing reports 5% or more of common stock, par value $0.0001 per share. Each figure below is the holder's own, as filed, with the filing linked.
Layoffs on record: 1 filing or notice, latest 2025-11-06 (layoff filings and notices).
Holders
| Holder | Percent | Shares | Latest | Event date |
|---|---|---|---|---|
| Aberdeen Group plc | 9.86% | 10,797,702 | SCHEDULE 13G/A, 2026-10-30 | 2026-09-30 |
| Telemark Asset Management, LLC | 7.3% | 7,846,411 | SCHEDULE 13G, 2026-07-29 | 2026-07-24 |
| 5.2% | 5,592,972 | SCHEDULE 13G/A, 2026-08-13 | 2026-06-30 | |
| Integrated Core Strategies (US) LLC | 5.2% | 4,734,627 | SCHEDULE 13G, 2026-04-14 | 2026-04-09 |
| Opaleye Management Inc. | 2.37% | 2,590,000 | SCHEDULE 13G/A, 2026-11-06 | 2026-09-30 |
| Fmr LLC | 0.6% | 584,367 | SCHEDULE 13G/A, 2025-11-05 | 2025-09-30 |
| Casdin Capital, LLC | 0% | 0 | SCHEDULE 13G/A, 2025-11-14 | 2025-09-30 |
| The Vanguard Group | 0% | 0 | SCHEDULE 13G/A, 2026-03-26 | 2026-03-13 |
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More on Codexis, Inc. and the records around it.
Codexis, Inc. elsewhere on the record
- Leadership changes Apr 2026
- Layoff notices Nov 2025
- Who owns it
Companies that share a holder
- Telemark Asset Management, LLC also filed stake reports on 2 other companies: Andersen Group Inc. (Aug 2026), Biodesix, Inc. (Jan 2026)
- Casdin Capital, LLC also filed stake reports on 3 other companies: Standard BioTools Inc. (Jun 2026), BIOLIFE SOLUTIONS INC (Jun 2026), Relay Therapeutics, Inc. (Feb 2026)
- Aberdeen Group plc also filed stake reports on 12 other companies: Equillium, Inc. (Nov 2026), IMMUNIC, INC (Nov 2026), Corbus Pharmaceuticals Holdings, Inc. (Oct 2026)
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What the Full Report on Codexis, Inc. covers
The Short Answer
Which Company This Is
What They Do, And How The Money Works
Who Runs It, And How To Reach Them
The Money: Funding, Valuation, Runway
Who Pays Them, And Who They Are Up Against
Warning Bells
Sources
19 cited sources found
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Voting and dispositive power, as filed
| Reporting person | Sole voting | Shared voting | Sole dispositive | Shared dispositive | Filing |
|---|---|---|---|---|---|
| Aberdeen Group plc | 0 | 10,797,702 | 0 | 10,797,702 | SCHEDULE 13G/A, 2026-10-30 |
| abrdn Inc. | 0 | 10,797,702 | 0 | 10,797,702 | SCHEDULE 13G/A, 2026-10-30 |
| Telemark Asset Management, LLC | 0 | 7,846,411 | 0 | 7,846,411 | SCHEDULE 13G, 2026-07-29 |
| Telemark Fund LP | 0 | 7,846,411 | 0 | 7,846,411 | SCHEDULE 13G, 2026-07-29 |
| Colin S. McNay | 0 | 7,846,411 | 0 | 7,846,411 | SCHEDULE 13G, 2026-07-29 |
| Lytton Laurence W | 5,498,680 | 94,292 | 5,498,680 | 94,292 | SCHEDULE 13G/A, 2026-08-13 |
| Opaleye Management Inc. | 0 | 2,590,000 | 0 | 2,590,000 | SCHEDULE 13G/A, 2026-11-06 |
| Opaleye, L.P. | 0 | 2,590,000 | 0 | 2,590,000 | SCHEDULE 13G/A, 2026-11-06 |
| James Silverman | 0 | 2,590,000 | 0 | 2,590,000 | SCHEDULE 13G/A, 2026-11-06 |
The latest filing identifies Codexis, Inc. by CUSIP 192005106 and gives its principal executive office as 200 Penobscot Drive, Redwood City, CA 94063 (SCHEDULE 13G/A, 2026-11-06).
What the percentages are based on
See response to Item 11 on each cover page. The percentage reported in this Statement is based upon 107,561,409 shares of Common Stock outstanding according to the final prospectus supplement filed by the Issuer with the U.S. Securities and Exchange Commission on July 24, 2026.Telemark Asset Management, LLC, SCHEDULE 13G, 2026-07-29
2.37 % Based on 109,505,765 Common Shares outstanding as of 08/06/2026 as reported by Codexis, Inc. on 08/11/2026, in its Form 10-Q Statement.Opaleye Management Inc., SCHEDULE 13G/A, 2026-11-06
Timeline
| Filed | Holder | Percent | Filing |
|---|---|---|---|
| 2025-07-30 | Aberdeen Group plc | 5.07% | SCHEDULE 13G |
| 2025-08-14 | Casdin Capital, LLC | 8.7% | SCHEDULE 13G/A |
| 2025-11-05 | Fmr LLC | 0.6% | SCHEDULE 13G/A |
| 2025-11-07 | Aberdeen Group plc | 4.53% | SCHEDULE 13G/A |
| 2025-11-14 | Casdin Capital, LLC | 0% | SCHEDULE 13G/A |
| 2025-12-05 | Opaleye Management Inc. | 9.95% | SCHEDULE 13G/A |
| 2026-01-15 | Aberdeen Group plc | 8.82% | SCHEDULE 13G/A |
| 2026-01-23 | Opaleye Management Inc. | 7.52% | SCHEDULE 13G/A |
| 2026-03-11 | Lytton Laurence W | 5% | SCHEDULE 13G |
| 2026-03-26 | The Vanguard Group | 0% | SCHEDULE 13G/A |
| 2026-04-14 | Integrated Core Strategies (US) LLC | 5.2% | SCHEDULE 13G |
| 2026-05-05 | Opaleye Management Inc. | 7.47% | SCHEDULE 13G/A |
| 2026-07-20 | Aberdeen Group plc | 7.45% | SCHEDULE 13G/A |
| 2026-07-29 | Telemark Asset Management, LLC | 7.3% | SCHEDULE 13G |
| 2026-08-13 | Lytton Laurence W | 5.2% | SCHEDULE 13G/A |
| 2026-10-30 | Aberdeen Group plc | 9.86% | SCHEDULE 13G/A |
| 2026-11-06 | Opaleye Management Inc. | 2.37% | SCHEDULE 13G/A |
Questions about who owns Codexis, Inc.
Who owns Codexis, Inc.?
On this page, 4 holders' latest ownership filings report 5% or more of Codexis, Inc.: Aberdeen Group plc (9.86%, Schedule 13G/A, filed 2026-10-30), Telemark Asset Management, LLC (7.3%, Schedule 13G, filed 2026-07-29), Lytton Laurence W (5.2%, Schedule 13G/A, filed 2026-08-13) and Integrated Core Strategies (US) LLC (5.2%, Schedule 13G, filed 2026-04-14).
Who are the largest shareholders of Codexis, Inc.?
By the share each holder's latest filing reports, the largest holders of Codexis, Inc. on this page are Aberdeen Group plc (9.86%), Telemark Asset Management, LLC (7.3%) and Lytton Laurence W (5.2%).
When did Aberdeen Group plc buy into Codexis, Inc.?
The filings give report dates, not purchase dates. The earliest filing from Aberdeen Group plc on this page is the Schedule 13G, filed 2025-07-30, reporting 5.07%; its latest, Schedule 13G/A filed 2026-10-30, reports 9.86%.
Is this an audit or a full list?
Neither. The ownership list on this page covers only Schedule 13D and 13G filings, which holders make once they own 5% or more of a class of shares (an amendment can report a stake that has since fallen below 5%); 17 such filings are shown here. It is not an audit and not a full cap table: holders who never reached 5% are not listed, and a stake can change after its latest filing.
Also on the record for Codexis, Inc.
| Date | Record | Figure | Our page | Source |
|---|---|---|---|---|
| April 14, 2026 | Leadership change: Dennis Wolf departs | — | Leadership changes | Source |
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Where this comes from
Anyone who comes to own more than 5% of a class of a listed company's voting shares has to tell the SEC. A holder who may seek to change or influence the company files Schedule 13D, and has to say in Item 4, "Purpose of Transaction", what it intends to do. A holder with no such intent, such as many index and passive funds, may file the shorter Schedule 13G. Both are amended when the stake changes, including when it falls below 5%.
Every row here is one of those filings, linked to the filing itself. Percentages and share counts are exactly as the holder filed them, for the holder the filing names; an amendment showing a lower figure is shown as filed. Item 4 text is quoted word for word. Where a 13D's Item 4 contains a sentence stating a definite intent to influence the company, that sentence is marked and quoted; the page does not describe the holder or its motives beyond the words it filed. Nothing here is investment advice.